【多选题】【消耗次数：1】
植物衰老的生理变化包括（）。
①
蛋白质显著下降
②
核酸含量降低
③
光合速率下降
④
呼吸速率下降
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由于蛋白质的互补作用，玉米蛋白质的生理价值是60%，小麦蛋白质的生理价值是67%，黄豆蛋白质的生理价值是65%，而它们综合在一起的生理价值是( )。
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